SCARABAEIDAE |
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Aphodiinae |
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Aphodiini |
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Aphodius abusus Fall (3,15) |
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A. bicolor Say (12,15) |
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A. cavidomus Brown (2,17,38) |
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A. coloradensis Horn (3,34) |
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A. concavus Say (3,15) |
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A. cynomysi Brown (11,17,38) |
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A. denticulatus Haldeman (34) |
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A. dentigerulus Brown (11,21,34) |
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A. distinctus (Muller) (3,12,15) |
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A. femoralis Say (3,12,15) |
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A. fimetarius (L.) (2,12,15,64) |
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A. granarius (L.) (3,12,15) |
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A. haemorrhoidalis (L.) (1) |
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A. haldemani Horn (18) |
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A. insolitus Brown (11,18) |
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A. lividus (Olivier) (2,3,12,15,63) |
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A. lutulentus Haldeman (3,12,15,35) |
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A. magnificans Robinson (11) |
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A. neodistinctus Brown (11,19) |
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A. oklahomensis Brown (11,18) |
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A. peculiosus Schmidt (15, as longitarsus Fall) |
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A. pseudabusus Cartwright (24) |
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A. pseudolividus Balthasar (47) |
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A. punctissimus Brown (11,18) |
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A. rubeolus Beauvois (2,3,12,15,46) |
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A. rusicola Melsheimer (2,3,12,15,46) |
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A. russeus Brown (11,18) |
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A. serval Say (1,15) |
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A. stercorosus Melsheimer (2,12,15) |
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A. tenuistriatus Horn (2,3,12,15,17,38) |
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A. terminalis Say (2,15) |
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A. vittatus Say (10,12,15) |
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A. walshii Horn (15) |
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Didactyliini |
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Aidophus parcus (Horn) (28)(? 59, as Dialytes) |
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Eupariini |
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Ataenius apicalis Hinton (1) |
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A. cognatus (LeConte) (12,15,25,46) |
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A. figurator Harold (2,3,25,37,46) |
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A. gracilis (Melsheimer) (2,3,25,46) |
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A. hesperius Cartwright (1,25,46) |
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A. imbricatus (Melsheimer) (2,12,25) |
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A. inquisitus Horn (46) |
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A. oklahomensis Brown (11,25) |
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A. platensis (Blanchard) (2,3,12,25,47) |
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A. puncticollis (LeConte) (2) |
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A. punctifrons Cartwright (25) |
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A. robustus Horn (3,12,25) |
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A. spretulus (Haldeman) (2,3,12,25,46) |
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A. strigatus (Say) (2,3,12,25,46) |
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A. texanus Harold (2,3,36)(38, as abditus texanus Harold) |
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(12, as abditus Haldeman) |
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A. wenzelii Horn (25) |
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Martineziella dutertrei
(Chalumeau) (1,46,62) |
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Pseudataenius socialis (Horn) (25) |
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Psammodiini |
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Platytomus atlanticus
Cartwright (1) |
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P. longulus (Cartwright) (1,47) |
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P. notialis Cartwright (1) |
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Psammodius quinqueplicatus Horn (15, sp. prob. not correct) |
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Rhyssemus neglectus Brown (36) |
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R. sonatus LeConte (15) |
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Scarabaeinae |
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Canthonini |
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Canthon chalcites (Haldeman) (2,12,15,38,50,63,64) |
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C. depressipennis LeConte (38) |
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C. ebenus (Say) (3,12,15,50,63) |
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C. hudsonias (Forster) (2)(15,38,39,50,63, as laevis Drury) |
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(12,46,64, as pilularius L.) |
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C. imitator Brown (1,12,22,54,63) |
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C. lecontei Harold (2,3,12,15,38,63) |
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C. praticola LeConte (3,12,15,54,63) |
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C. probus (Germar) (54,64) |
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C. vigilans LeConte (2,12,15,54,63,64) |
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C. viridis (Beauvois) (1,12,15,46,63,64) |
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Deltochilum gibbosus (Fab.) (3,63) |
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Melanocanthon nigricornis (Say) (3)(15,47,63, as Canthon) |
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Pseudocanthon perplexus
(LeConte) (2,12,64)(46,63, as Canthon) |
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Coprini |
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Copris f. fricator (Fab.) (2,3,12,49)(15,39,63, as tullius Olivier) |
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C. minutus (Drury) (2,12,15,49,63,64) |
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C. r. remotus LeConte (12,49) |
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Dichotomiini |
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Ateuchus histeroides Weber (3,46,62)(15,50,53, as Choeridium) |
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A. lecontei Harold (10,63 - id ?) |
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Dichotomius carolinus (L.) (3,12,63,64)(15,39, as Pinotus) |
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Oniticellini |
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Euoniticellus intermedius
(Reiche) (66) |
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Onthophagini |
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Onthophagus cavernicollis Howden
& Cartwright (43,63) |
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O. cynomisi Brown (11,16,38,43,63) |
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O. gazella Fab. (1,12,44,46) |
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O. h. hecate (Panzer) (2,12,14,15,39,43,46,50,63) |
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O. landolti texanus Schaeffer (12,43) |
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O. medorensis Brown (3,43,63)(14,15,39, as guatemalensis Bates) |
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O. oklahomensis Brown (1,12,16,38,43,63) |
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O. o. orpheus (Panzer) (1,12,14,15,39,43,46,63) |
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O. pennsylvanicus Harold (3,12,14,15,38,39,43,47,63) |
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O. s. striatulus (Beauvois) (15,43) |
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O. subaeneus (Beauvois) (43,63)(14,15,39, as cribricollis Horn) |
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O. tuberculifrons Harold (3,12,14,15,39,43,63) |
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O. velutinus Horn (1,47) |
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Phanaeini |
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Phanaeus difformis LeConte (4,12,13,15,29,63) |
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P. t. triangularis (Say) (12,29) |
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P. vindex MacLeay (4,12,13,29,46,63,64)(15,39,50, as carnifex
L.) |
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Melolonthinae |
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Sericini |
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Serica aspera Dawson ? (1) |
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S. atratula atratula LeConte (3,26,27) |
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S. atratula monita Dawson (11) |
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S. campestris Dawson (27,47) |
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S. contorta Dawson (11,26,27) |
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S. intermixta Blatchley (3) |
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S. mystaca Dawson (27) |
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S. ochrosoma Dawson (27) |
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S. parallela Casey (1) |
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S. sericea (Illiger) (3,27) |
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Melolonthini |
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Hypothyce mixta Howden ? (1) |
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Phyllophaga aemula (Horn) (7) |
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P. affabilis (Horn) (3) |
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P. anxia (LeConte) (7,8) |
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P. arkansana (Schaeffer) (7) |
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P. aurea Luginbill &
Painter (11) |
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P. bipartita (Horn) (7,8) |
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P. calceata (LeConte) (7,8,47) |
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P. congrua (LeConte) (7,8) |
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P. corrosa (LeConte) (5,7,39) |
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P. crassissima (Blanchard) (3,7,8) |
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P. crenulata (Froelich) (3,7,8,46,50) |
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P. cribrosa (LeConte) (3,8,32,46) |
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P. crinita (Burmeister) (3,46) |
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P. curialis Reinhard (1) |
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P. delata (Horn) (8) |
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P. dentex (Bates) (48) |
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P. ephilida (Say) (3,8) |
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P. farcta (LeConte) (1) |
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P. fervida (Fab.) (1) |
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P. forbesi Glasgow (7,46) |
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P. forsteri (Burmeister) (8) |
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P. fraterna Harris (3,8) |
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P. futilis (LeConte) (5,8,47) |
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P. glabricula (LeConte) (3,5) |
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P. gracilis (Burmeister) (3) |
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P. hirticula (Knoch) (48) |
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P. hirtiventris (Horn) (7,39) |
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P. hornii (Smith) (7) |
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P. ilicis (Knoch) (7,8,52) |
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P. implicita (Horn) (7) |
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P. incuria Sanderson (48) |
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P. inepta (Horn) (7) |
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P. koehleriana Saylor (3) |
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P. lanceolata (Say) (5,30,46) |
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P. longitarsa (Say) (3,39) |
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P. luctuosa (Horn) (48) |
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P. macmurryi Saylor (11,55) |
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P. marginalis (LeConte) (8) |
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P. micans (Knoch) (8) |
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P. pearliae Davis (8) |
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P. praetermissa (Horn) (7) |
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P. profunda (Blanchard) (7) |
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P. prunina (LeConte) (8) |
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P. pulcher (Linell) (8) |
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P. quercus (Knoch) (3,5,7,50) |
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P. rubiginosa (LeConte) (7,46) |
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P. rugosa (Melsheimer) (8) |
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P. scitula (Horn) (48) |
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P. sodalis Reinhard (52) |
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P. squamipilosa Saylor (52) |
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P. submucida (LeConte) (7,8,46) |
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P. sylvatica Sanderson (8) |
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P. torta (LeConte) (46,48) |
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P. tristis (Fab.) (8) |
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P. vanalleri (Schaeffer) (8) |
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P. vehemens (Horn) (1) |
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Polyphylla decimlineata (Say) (9) |
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P. hammondi LeConte (9,46)(50, as oklahomensis Hatch) |
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Diplotaxini |
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Diplotaxis angularis LeConte (61) |
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D. atlantis Fall (61) |
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D. belfragei Fall (3,61) |
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D. blanchardi Vaurie (1,61) |
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D. frondicola (Say) (3,46,61) |
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D. harperi Blanchard (3,46,61) |
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D. haydenii LeConte (1) |
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D. lengii Fall (11,39,61) |
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D. maura Fall (46) |
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D. punctata LeConte (3,61) |
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D. punctatorugosa Blanchard (61) |
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D. rudis (LeConte) (3,60) |
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D. subangulata LeConte (3) |
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D. thoracica Fall (1,61) |
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D. truncatula LeConte (46) |
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Macrodactylini |
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Dichelonyx linearis (Gyllenhal) (22)(57, as elongata Fab.) |
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D. subvittata (LeConte) (1,10) |
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Macrodactylus angustatus
(Beauvois) (38) |
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M. subspinosus (Fab.) (3,39,55) |
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M. uniformis Horn (3) |
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Rutelinae |
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Anomalini |
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Anomala binotata (Gyllenhal) (3,37,47,51) |
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A. cavifrons LeConte (3,37,51) |
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A. f. flavipennis Burmeister (3,37,46,51) |
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A. flavipennis luteipennis
LeConte (3,37,51) |
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A. innuba (Fab.) (3,37,38,46,51) |
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A. ludoviciana Schaeffer (3,37,51) |
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A. marginata (Fab.) (3,46)(38, as Spilota) |
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A. undulata Melsheimer (3) |
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Popillia japonica Newman (1) |
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Strigoderma arboricola (Fab.) (3,46) |
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S. knausi (Brown) (37, as pygmaea (Fab.))(11, as oklahomensis
Hatch) |
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(38, as Strigodermella oklahomensis
Hatch) |
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Rutelini |
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Cotalpa lanigera (L.) (3) |
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C. subcribrata Wickham (3,65) |
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Parastasia brevipes (LeConte) (3,8,46) |
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Pelidnota punctata (L.) (3,37,46)(38,39,50, as lutea texensis
Casey) |
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Dynastinae |
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Cyclocephalini |
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Cyclocephala borealis Arrow (10) |
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C. hirta LeConte (3,37,46) |
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C. immaculata (Olivier) (3,37,50,58)(38,39, as Ochrosidia rufifrons
Casey) |
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C. longula LeConte (3) |
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C. melanocephala (Fab.) (3,46) |
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C. pasadenae (Casey) (3) |
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C. robusta LeConte (3)(37, as C. nigricollis Burmeister) |
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Dyscinetus morator (Fab.) (3,46)(50, as trachypygus Burmeister) |
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D. picipes (Burmeister) (3) |
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Pentodontini |
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Aphonus texanus Gill &
Howden (1,37) |
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Euetheola humilis rugiceps
(LeConte) (3,37,46) |
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Tomarus gibbosus gibbosus
(DeGeer) (3,37)(23,31, as Bothynus) |
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(46, as Ligyrus) |
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T. relictus (Say) (3,37)(23, as Bothynus) |
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Oryctini |
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Strategus aloeus (L.) (3) |
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S. antaeus (Drury) (3,37) |
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S. mormon Burmeister (3,37) |
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Xyloryctes jamaicensis (Drury) (3)(39, as satyrus Fab.) |
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Phileurini |
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Phileurus truncatus (Beauvois) (1) |
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P. valgus (L.) (3,46) |
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Dynastini |
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Dynastes tityus tityus (L.) (3,33) |
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Cetoniinae |
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Gymnetini |
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Cotinis nitida (L.) (3,32,38,46,50) |
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Cetoniini |
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Euphoria fulgida (Fab.) (3,32) |
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E. herbacea occidentalis Knaus (50) |
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E. inda (L.) (3) |
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E. kerni Haldeman (3,38,39,46)38,39, as E. clarki LeConte) |
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(39, as E. texanus Schaufuss) |
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E. sepulchralis (Fab.) (3,38,39,46) |
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Stephanucha pilipennis Kraatz (3) |
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Cremastocheilini |
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Cremastocheilus knochi LeConte (3) |
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C. retractus LeConte (1) |
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Trichiini |
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Osmoderma eremicola (Knoch) (3,41)(42, as subplanata Casey) |
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Trichiotinus lunulatus (Fab.) (1) |
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T. piger (Fab.) (6,38,40,42) |
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T. texanus (Horn) (6,40,42,46) |
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Trigonopeltastes delta
(Forster) (3,38,42,46,50) |
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Valgini |
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Valgus canaliculatus (Olivier) (1,45) |
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V. seticollis (Beauvois) (1) |
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1. In OSU Collection - Det. D. C. Arnold |
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2. In OSU Collection - Det. W. J. Brown |
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3. In OSU Collection - Det. O. L. Cartwright |
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4. In OSU Collection - Det. W. D. Edmonds |
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5. In OSU Collection - Det. A. A. Granovsky |
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6. In OSU Collection - Det. C. E. Hoffmann |
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7. In OSU Collection - Det. P. Luginbill |
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8. In OSU Collection - Det. M. W. Sanderson |
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9. In OSU Collection - Det. R. M. Young |
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10. In OSU
Collection - Determiner unknown |
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11.
Reported in Coleoptera Catalog |
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12.
Reported by Blume, 1985 |
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13.
Reported by Blume & Aga, 1978 |
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14.
Reported by Brown, 1925 |
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15.
Reported by Brown, 1927a |
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16.
Reported by Brown, 1927b |
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17.
Reported by Brown, 1927c |
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18.
Reported by Brown, 1928a |
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19.
Reported by Brown, 1928b |
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20.
Reported by Brown, 1929a |
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21.
Reported by Brown, 1929b |
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22.
Reported by Brown, 1946 |
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23.
Reported by Cartwright, 1959 |
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24.
Reported by Cartwright, 1972 |
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25.
Reported by Cartwright, 1974 |
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26.
Reported by Dawson, 1947 |
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27.
Reported by Dawson, 1952 |
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28.
Reported by Dellacasa et al., 2001 |
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29.
Reported by Edmonds, 1994 |
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30.
Reported by Fenton, 1937 |
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31.
Reported by Fenton, 1938 |
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32.
Reported by Fenton, 1940 |
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33.
Reported by Fenton, 1941 |
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34.
Reported by Gordon, 1976 |
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35.
Reported by Gordon, 1983 |
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36.
Reported by Gordon & Cartwright, 1980 |
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37.
Reported by Hardy, 1991 |
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38.
Reported by Hatch, 1930 |
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39.
Reported by Hatch & Ortenburger, 1930 |
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40.
Reported by Hoffmann, 1935 |
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41.
Reported by Hoffmann, 1939 |
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42.
Reported by Howden, 1968 |
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43.
Reported by Howden & Cartwright, 1963 |
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44.
Reported by Hunter & Fincher, 1985 |
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45.
Reported by Jameson & Swoboda, 2005 |
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46.
Reported by Kondratieff et al., 2005 |
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47.
Reported by Kondratieff et al., 2007 |
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48.
Reported by Luginbill & Painter, 1953 |
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49.
Reported by Matthews, 1961 |
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50.
Reported by Ortenburger & Hatch, 1926 |
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51.
Reported by Potts, 1977 |
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52.
Reported by Riley & Wolfe, 2003 |
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53.
Reported by Robinson, 1948a |
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54.
Reported by Robinson, 1948b |
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55.
Reported by Sanborn, 1912 |
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56.
Reported by Saylor, 1940 |
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57.
Reported by Saylor, 1945a |
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58.
Reported by Saylor, 1945b |
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59.
Reported by Skelley & Gordon, 2002 |
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60.
Reported by Vaurie, 1958 |
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61.
Reported by Vaurie, 1960 |
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62.
Reported by Vogt et al., 2001 |
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63.
Reported by Wallace & Drew, 1965 |
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64.
Reported by Woodruff, 1973 |
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65.
Reported by Young, 2005 |
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66. In OSU
Collection - Det. R. A. Grantham |
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48 genera -
236 species & subspecies |
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